After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died. Gramma was born in her native habitat and was estimated to be about 141 years old, zoo officials said. She died on November 20. It’s not clear exactly when the tortoise arrived at the San Diego Zoo, but zoo officials said she came from the Bronx Zoo in either 1928 or 1931 as part of their first group of Galapagos tortoises.

Ding, J. (2025, November 30). Gramma, the Galapagos tortoise of San Diego Zoo, dies at about 141. BusinessMirror, p. A7.